Picea taiga woodland

This habitat comprises of mesic to herb-rich forest vegetation on mineral soils in the boreal and boreonemoral zones. The soils are often podzolic, but other soil types occur and the humus is raw or mull. The canopy is often dominated by Picea abies but, despite the name of the habitat, stands dominated by Pinus sylvestris or Betula pendula or mixtures of these trees are also common. Other tree species include Alnus incana, Betula pubescens, Populus tremula, Salix caprea, Sorbus aucuparia, and in the boreonemoral and southern boreal subzones also Acer platanoides, Alnus glutinosa, Quercus robur, Tilia cordata, Ulmus glabra and Ulmus laevis. Under natural conditions, forest succession will lead to the development of a Picea abies forest, but the proportions of tree species are nowadays largely regulated by forestry. The shrub layer is best developed in moist herb-rich stands, where Frangula alnus, Lonicera xylosteum, Prunus padus, Ribes spp., Rubus idaeus and other shrubs can form dense thickets. On mesic sites, by contrast, the only true shrubs are Juniperus communis, Salix caprea and other Salix spp. Understorey vegetation varies from the dwarf shrub and feather moss dominated vegetation in mesic situations to the most luxurious and species-rich herb dominated vegetation. In mesic situations, Vaccinium myrtillus usually dominates, followed by Vaccinium vitis-idaea, Linnaea borealis and, in the middle and northern boreal subzones, even Empetrum nigrum, Ledum palustre and Vaccinium uliginosum. The commonest herbs are Convallaria majalis, Dryopteris carthusiana, Epilobium angustifolium, Maianthemum bifolium, Melampyrum pratense, Melampyrum sylvaticum, Pteridium aquilinum, Solidago virgaurea and Trientalis europaea, while species like Lathyrus vernus and Oxalis acetosella grow on slightly more nutrient-rich soils. Calamagrostis arundinacea and Deschampsia flexuosa are the most abundant grasses on mesic sites. Common graminoids also include Carex digitata, Carex globularis, Deschampsia cespitosa, Luzula pilosa and Melica nutans. There is great compositional variation in the herb layer of the richer sites, depending on the geographic location, soil moisture, soil nutrient status and canopy composition. In general, the number of herb species is high, and there are also many graminoids, but dwarf shrubs are few or non-existent. However, the number of vascular species decreases towards the north of the range. In addition to species thriving on mesic sites, examples of typical herb species are Aegopodium podagraria, Anemone nemorosa, Angelica sylvestris, Anthriscus sylvestris, Athyrium filix-femina, Cirsium helenioides, Cornus suecica, Corydalis solida, Dryopteris carthusiana, Dryopteris expansa, Filipendula ulmaria, Fragaria vesca, Galium boreale, Geranium sylvaticum, Geum rivale, Gymnocarpium dryopteris, Hepatica nobilis, Matteuccia struthiopteris, Paris quadrifolia, Pulmonaria obscura, Ranunculus auricomus, Ranunculus fallax, Stellaria nemorum, and common grasses are Agrostis capillaris, Calamagrostis purpurea, Melica nutans, Milium effusum and Poa nemoralis. On mesic sites the moss layer is usually continuous and dominated by feather mosses like Pleurozium schreberi and Hylocomium splendens. Other common species are Dicranum fuscescens, D. majus, D. polysetum, D. scoparium, Polytrichum commune, Ptilium crista-castrensis and on slightly more nutrient-rich sites Climacium dendroides, Rhodobryum roseum and Rhytidiadelphus triquetrus. The abundance and number of liverwort species, like Barbilophozia lycopodioides, increases towards north. On mesic sites there may even be some terricolous lichens. On herb-rich sites, the cover of the moss layer is usually small, and feather mosses are scarce. Instead, there is a rich flora of nutrient-demanding mosses and liverworts like Brachythecium spp., Cirriphyllum piliferum, Plagiochila asplenioides, Plagiomnium spp., Plagiothecium spp., Pseudobryum cinclidioides and Rhizomnium spp. After a major disturbance such as windfall, forest fire or regeneration cutting, herbs and grasses increase, Vaccinium myrtillus declines and bryophytes decrease.

Indicators of good quality:

• Natural composition of canopy

• Structural diversity/ complexity with (semi)natural age structure or completeness of layers

• Typical flora and fauna composition of the region

• Presence of old trees and a variety of dead wood (lying or standing) and the associated flora, fauna and fungi

• Presence of natural disturbance such as treefall openings with natural regeneration

• Long historical continuity (ancient woodland) with high species diversity

• Survival of larger stands of forest without anthropogenic fragmentation and isolation (to support fauna which need large undisturbed forests)

• Absence of non-native species in all layers (flora & fauna)

• No signs of eutrophication or pollution

• No man-induced very high population levels of ungulates

Synthesis of Red List assessment

Confidence in the assessment

Pressures and threats